Editing LDAP connector with more than 1 group in the Default Group Memberships will have error

Steps to reproduce

1. Create a LDAP connector at Administration >> User Directories
2. Set LDAP Permissions to "Read Only, with Local Groups"
3. Insert "jira-users,jira-developers" to Default Group Memberships
4. Test the setting and save
5. Click 'Edit' link for the same LDAP connector and test the setting will has an error (see screenshot: default_group.png). Default Group Memberships showing "jira-users|jira-developers"

Editing delegated LDAP with the same setting does not have the error.