JIRA issues macro displays "USER_UNKNOWN"

com.atlassian.confluence.extra.jira.TrustedAppsException: USER_UNKNOWN;	Unknown User: {0};	["abc@abc.com"]
	at com.atlassian.confluence.extra.jira.DefaultJiraIssuesManager$1.handle(DefaultJiraIssuesManager.java:112)

Steps to reproduce:

1. Set up JIRA and Confluence on Trusted application
2. Set up a Crowd instance connecting to JIRA and Confluence.
3. The username for the user in Confluence is abc@abc.com
4. In Crowd, create an alias for this user for JIRA as 'test'
5. So, the username for the same user in JIRA is test
6. Now login to JIRA with this user and then create an issue.
7. In Confluence, create a page with JIRA issues macro and insert a link to the issue created above in JIRA.
8. The error message above should show up..

JIRA issues macro is not able to translate the user 'test' to 'abc@abc.com'

You can also find more information in JIRA issues macro displays "USER_UNKNOWN".