Stuck mail queue

The mail queue gets stuck (no more mails sent) and can't be flushed.
Note that we did an export/import since the upgrade to 2.6.

This seems to be the corresponding error in the logs based on the first stuck mail date (there is no corresponding error i the mail server logs):

2004-03-22 12:14:18,622 ERROR [atlassian.jira.workflow.OSWorkflowManager] An exception occurred
com.opensymphony.workflow.InvalidActionException: Action 1 is invalid
at com.opensymphony.workflow.AbstractWorkflow.doAction(AbstractWorkflow.java:480)
at com.atlassian.jira.workflow.OSWorkflowManager.doWorkflowAction(OSWorkflowManager.java:150)
at com.atlassian.jira.web.action.issue.ResolveIssue.doExecute(ResolveIssue.java:116)
at webwork.action.ActionSupport.execute(ActionSupport.java:151)
at com.atlassian.jira.action.JiraActionSupport.execute(JiraActionSupport.java:49)
at webwork.dispatcher.GenericDispatcher.executeAction(GenericDispatcher.java:131)
at com.atlassian.jira.web.dispatcher.JiraServletDispatcher.service(JiraServletDispatcher.java:186)
at javax.servlet.http.HttpServlet.service(HttpServlet.java:853)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:247)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.opensymphony.module.sitemesh.filter.PageFilter.parsePage(PageFilter.java:129)
at com.opensymphony.module.sitemesh.filter.PageFilter.doFilter(PageFilter.java:61)
at com.atlassian.jira.web.filters.SitemeshExcludePathFilter.doFilter(SitemeshExcludePathFilter.java:36)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.seraph.filter.SecurityFilter.doFilter(SecurityFilter.java:164)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.seraph.filter.LoginFilter.doFilter(LoginFilter.java:181)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.util.profiling.filters.ProfilingFilter.doFilter(ProfilingFilter.java:132)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.jira.web.filters.ActionCleanupDelayFilter.doFilter(ActionCleanupDelayFilter.java:37)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.johnson.filters.JohnsonFilter.doFilter(JohnsonFilter.java:96)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.jira.web.filters.gzip.GzipFilter.doFilter(GzipFilter.java:60)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at com.atlassian.core.filters.AbstractEncodingFilter.doFilter(AbstractEncodingFilter.java:38)
at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:213)
at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:193)
at org.apache.catalina.core.StandardWrapperValve.invoke(StandardWrapperValve.java:256)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:643)
at org.apache.catalina.core.StandardPipeline.invoke(StandardPipeline.java:480)
at org.apache.catalina.core.ContainerBase.invoke(ContainerBase.java:995)
at org.apache.catalina.core.StandardContextValve.invoke(StandardContextValve.java:191)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:643)
at org.apache.catalina.core.StandardPipeline.invoke(StandardPipeline.java:480)
at org.apache.catalina.core.ContainerBase.invoke(ContainerBase.java:995)
at org.apache.catalina.core.StandardContext.invoke(StandardContext.java:2415)
at org.apache.catalina.core.StandardHostValve.invoke(StandardHostValve.java:180)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:643)
at org.apache.catalina.valves.ErrorDispatcherValve.invoke(ErrorDispatcherValve.java:171)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:641)
at org.apache.catalina.valves.ErrorReportValve.invoke(ErrorReportValve.java:172)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:641)
at org.apache.catalina.valves.AccessLogValve.invoke(AccessLogValve.java:509)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:641)
at org.apache.catalina.core.StandardPipeline.invoke(StandardPipeline.java:480)
at org.apache.catalina.core.ContainerBase.invoke(ContainerBase.java:995)
at org.apache.catalina.core.StandardEngineValve.invoke(StandardEngineValve.java:174)
at org.apache.catalina.core.StandardPipeline$StandardPipelineValveContext.invokeNext(StandardPipeline.java:643)
at org.apache.catalina.core.StandardPipeline.invoke(StandardPipeline.java:480)
at org.apache.catalina.core.ContainerBase.invoke(ContainerBase.java:995)
at org.apache.ajp.tomcat4.Ajp13Processor.process(Ajp13Processor.java:466)
at org.apache.ajp.tomcat4.Ajp13Processor.run(Ajp13Processor.java:585)
at java.lang.Thread.run(Thread.java:534)